The film starts as prequel, showing a small part of Alex's early life, including his capture by hunters. It soon moves to shortly after the point where the original left off, with the animals deciding to return to New York. They board an airplane on Madagascar, but crash-land in Africa, where each of the central characters meets others of the same species; Alex is reunited with his parents. Problems arise, and their resolution occupies much of the remainder of the film.

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